Flanker Mark Wilson has today been selected on the bench for England's Rugby World Cup clash with the USA.

Wilson, who began his career in Penrith, is named as a substitute after his recent struggle with a knee injury which ruled him out of England's opening 35-3 success over Tonga on Sunday.

Joe Cokanasiga, meanwhile, will provide firepower to England's backline in the absence of Manu Tuilagi after being picked on the left wing for Thursday's contest (11.45am kick-off).

Cokanasiga replaces Jonny May in one of 10 changes to the starting XV that launched their Pool C campaign in Sapporo.

Cokanasiga's presence makes up for Tuilagi being omitted from the 23 after the Leicester wrecking ball laid waste to Tonga with a devastating two-try salvo and series of marauding runs.

Captain Owen Farrell drops to the bench so George Ford, who leads the team, continues at fly-half inside a new-look centre combination of Piers Francis and Jonathan Joseph.

Ruaridh McConnochie wins his second cap on the right wing, with Elliot Daly continuing at full-back and Willi Heinz getting the nod ahead of Ben Youngs at scrum-half.

There are three survivors from the pack that start against Tonga in prop Joe Marler, flanker Tom Curry and number eight Billy Vunipola.

Coach Eddie Jones said: "The four-day turnaround is something we experimented with against Japan last autumn and so the players have had some experience of it.

"You have to make sure the two days before the game are exactly the same as you would normally have for any other game.

"What I see now is players handling it really well, but the game on Thursday will be the litmus test.

"It's our best 23 and we have looked at some players that would benefit not playing in this game, so have kept them out of the 23.

"But it's a great opportunity for another set of players to do the team proud.

"USA are a tough, physical team who are extremely well coached by Gary Gold. There are a number of players who our guys know really well and are a team we respect."

Kendal-born Wilson, 29, has so far won 15 international caps.